About the Authors

Keith M. Callenberg

Affiliation Carnegie Mellon-University of Pittsburgh Program in Computational Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Om P. Choudhary

Affiliation Carnegie Mellon-University of Pittsburgh Program in Computational Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Gabriel L. de Forest

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

David W. Gohara

Affiliation The Edward A. Doisy Department of Biochemistry and Molecular Biology, Saint Louis University, St. Louis, Missouri, United States of America

Nathan A. Baker

Affiliation Pacific Northwest National Laboratory, Richland, Washington, United States of America

Michael Grabe

mdgrabe@pitt.edu

Affiliations Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Computational and Systems Biology, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MG. Performed the experiments: KMC OPC. Analyzed the data: KMC. Contributed reagents/materials/analysis tools: KMC OPC GLdF DWG NAB MG. Wrote the paper: KMC OPC NAB MG.