About the Authors

John C. Morgan

Affiliation Liverpool School of Tropical Medicine, Liverpool, United Kingdom

Helen Irving

Affiliation Liverpool School of Tropical Medicine, Liverpool, United Kingdom

Loyce M. Okedi

Affiliation National Livestock Resources Research Institute, Tororo, Uganda

Andrew Steven

Affiliation Liverpool School of Tropical Medicine, Liverpool, United Kingdom

Charles S. Wondji

c.s.wondji@liverpool.ac.uk

Affiliation Liverpool School of Tropical Medicine, Liverpool, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CSW. Performed the experiments: JCM HI AS CSW. Analyzed the data: CSW. Contributed reagents/materials/analysis tools: LMO. Wrote the paper: CSW.