About the Authors

Dion Lepp

Contributed equally to this work with: Dion Lepp, Bryan Roxas

Affiliations Department of Pathobiology, University of Guelph, Guelph, Ontario, Canada, Guelph Food Research Centre, Agriculture and Agri-Food Canada, Guelph, Ontario, Canada

Bryan Roxas

Contributed equally to this work with: Dion Lepp, Bryan Roxas

Affiliation Department of Veterinary Science and Microbiology, University of Arizona, Tucson, Arizona, United States of America

Valeria R. Parreira

Affiliation Department of Pathobiology, University of Guelph, Guelph, Ontario, Canada

Pradeep R. Marri

Affiliation BIO5 Institute, University of Arizona, Tucson, Arizona, United States of America

Everett L. Rosey

Affiliation Veterinary Medicine Research and Development, Pfizer Animal Health, Kalamazoo, Michigan, United States of America

Joshua Gong

Affiliation Guelph Food Research Centre, Agriculture and Agri-Food Canada, Guelph, Ontario, Canada

J. Glenn Songer

Affiliation Department of Veterinary Science and Microbiology, University of Arizona, Tucson, Arizona, United States of America

Gayatri Vedantam

Affiliations Department of Veterinary Science and Microbiology, University of Arizona, Tucson, Arizona, United States of America, Southern Arizona VA Healthcare System, Tucson, Arizona, United States of America

John F. Prescott

prescott@uoguelph.ca

Affiliation Department of Pathobiology, University of Guelph, Guelph, Ontario, Canada

Competing Interests

One of the several funders (Pfizer; E. L. Rosey) had a role in the study design, and in the decision to publish. The authors do not believe that this competing interest affects the objectivity, analyses, interpretations or validity of the study. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Performed the experiments: DL BR VRP PRM. Analyzed the data: DL BR VRP. Contributed reagents/materials/analysis tools: JG JGS JFP. Wrote the paper: DL BR VRP JGS GV JFP. Conceived the research: JFP DL ELR JG JGS. Designed the experiments: DL BR VRP.