About the Authors

Yuyan Zhang

Affiliations Institute of Oceanology, Chinese Academy of Science, Qingdao, People's Republic of China, Graduate School of Chinese Academy of Sciences, Beijing, People's Republic of China

Shaoli Yang

Affiliation Institute of Oceanology, Chinese Academy of Science, Qingdao, People's Republic of China

Ming Liu

Affiliation Institute of Oceanology, Chinese Academy of Science, Qingdao, People's Republic of China

Chunxia Song

Affiliation Institute of Oceanology, Chinese Academy of Science, Qingdao, People's Republic of China

Ning Wu

Affiliations Institute of Oceanology, Chinese Academy of Science, Qingdao, People's Republic of China, Graduate School of Chinese Academy of Sciences, Beijing, People's Republic of China

Peixue Ling

Affiliation Institute of Biopharmaceuticals of Shandong Province, Jinan, People's Republic of China

Edward Chu

Affiliation Yale Cancer Center, Yale University School of Medicine, New Haven, Connecticut, United States of America

Xiukun Lin

linxiukun@yahoo.com

Affiliation Institute of Oceanology, Chinese Academy of Science, Qingdao, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EC XL. Performed the experiments: YZ CS NW. Analyzed the data: YZ SY NW ML EC XL. Contributed reagents/materials/analysis tools: PL. Wrote the paper: SY.