About the Authors

Shahan Derkarabetian

Affiliation Department of Biology, San Diego State University, San Diego, California, United States of America

David B. Steinmann

Affiliation Department of Zoology, Denver Museum of Nature and Science, Denver, Colorado, United States of America

Marshal Hedin

mhedin@sciences.sdsu.edu

Affiliation Department of Biology, San Diego State University, San Diego, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SD DBS MH. Performed the experiments: SD. Analyzed the data: SD MH. Contributed reagents/materials/analysis tools: SD DBS MH. Wrote the paper: SD MH.