About the Authors

Mark B. Bain

Mark.Bain@Cornell.edu

Affiliation Department of Natural Resources, Cornell University, Ithaca, New York, United States of America

Emily R. Cornwell

Affiliation Department of Microbiology and Immunology, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Kristine M. Hope

Affiliation Department of Microbiology and Immunology, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Geofrey E. Eckerlin

Affiliation Department of Natural Resources, Cornell University, Ithaca, New York, United States of America

Rufina N. Casey

Affiliation Department of Microbiology and Immunology, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Geoffrey H. Groocock

Affiliation Department of Microbiology and Immunology, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Rodman G. Getchell

Affiliation Department of Microbiology and Immunology, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Paul R. Bowser

Affiliation Department of Microbiology and Immunology, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

James R. Winton

Affiliation Western Fisheries Research Center, U.S. Geological Survey, Seattle, Washington, United States of America

William N. Batts

Affiliation Western Fisheries Research Center, U.S. Geological Survey, Seattle, Washington, United States of America

Allegra Cangelosi

Affiliation Northeast-Midwest Institute, Washington, D. C., United States of America

James W. Casey

Affiliation Department of Microbiology and Immunology, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MBB PRB JRW JC. Performed the experiments: ERC KMH GEE RNC GHG RGG WNB JC. Analyzed the data: MBB ERC KMH GEE WNB. Contributed reagents/materials/analysis tools: ERC KMH GEE RNC RGG WNB. Wrote the paper: MBB GHG PRB JRW AC JC. Provided funding and general research direction: AC.