About the Authors

Kiyotaka Tokuraku

Affiliations Department of Pharmacology and Experimental Neuroscience, University of Nebraska Medical Center, Omaha, Nebraska, United States of America, Department of Chemical Science and Engineering, Miyakonojo National College of Technology, Miyakonojo, Japan

Meg Marquardt

Affiliation Department of Pharmacology and Experimental Neuroscience, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Tsuneya Ikezu

tikezu@unmc.edu

Affiliation Department of Pharmacology and Experimental Neuroscience, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KT TI. Performed the experiments: KT MM. Analyzed the data: KT MM. Wrote the paper: KT TI.