About the Authors

Qunzhou Zhang

Affiliation Center for Craniofacial Molecular Biology, School of Dentistry, University of Southern California, Los Angeles, California, United States of America

Takayoshi Yamaza

Affiliation Center for Craniofacial Molecular Biology, School of Dentistry, University of Southern California, Los Angeles, California, United States of America

A. Paul Kelly

Affiliation Department of Dermatology, King-Harbor Medical Center, Los Angeles, California, United States of America

Shihong Shi

Affiliation Center for Craniofacial Molecular Biology, School of Dentistry, University of Southern California, Los Angeles, California, United States of America

Songlin Wang

Affiliation Salivary Gland Disease Center and the Molecular Laboratory for Gene Therapy, Capital Medical University School of Stomatology, Beijing, China

Jimmy Brown

Affiliation Department of Otolaryngology, Head & Neck Surgery, Stanford University School of Medicine, Stanford, California, United States of America

Lina Wang

Affiliation Department of Pathology, Kenneth Norris Jr. Cancer Center, University of Southern California Keck School of Medicine, Los Angeles, California, United States of America

Samuel W. French

Affiliation Department of Pathology, Harbor-UCLA Medical Center, Torrance, California, United States of America

Songtao Shi

anhle@usc.edu (ADL); songtaos@usc.edu (SS)

Affiliation Center for Craniofacial Molecular Biology, School of Dentistry, University of Southern California, Los Angeles, California, United States of America

Anh D. Le

anhle@usc.edu (ADL); songtaos@usc.edu (SS)

Affiliation Center for Craniofacial Molecular Biology, School of Dentistry, University of Southern California, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: QZ TY SS AL. Performed the experiments: QZ TY SS LW. Analyzed the data: QZ TY SWF SS AL. Contributed reagents/materials/analysis tools: APK SW JB. Wrote the paper: QZ SS AL. Provide patient samples: AK.