About the Authors

Paige S. Davies

Contributed equally to this work with: Paige S. Davies, Anne E. Powell

Affiliation Department of Dermatology, Knight Cancer Institute, Oregon Stem Cell Center, Oregon Health & Science University, Portland, Oregon, United States of America

Anne E. Powell

Contributed equally to this work with: Paige S. Davies, Anne E. Powell

Affiliation Department of Cell and Developmental Biology, Oregon Health & Science University, Portland, Oregon, United States of America

John R. Swain

Affiliation Department of Dermatology, Knight Cancer Institute, Oregon Stem Cell Center, Oregon Health & Science University, Portland, Oregon, United States of America

Melissa H. Wong

wongme@ohsu.edu

Affiliations Department of Dermatology, Knight Cancer Institute, Oregon Stem Cell Center, Oregon Health & Science University, Portland, Oregon, United States of America, Department of Cell and Developmental Biology, Oregon Health & Science University, Portland, Oregon, United States of America

Competing Interests

The authors declare commercial funding from Proctor & Gamble Pharmaceuticals and confirm the receipt of this funding does not alter their adherence to all PLoS ONE policies regarding sharing data and materials.

Author Contributions

Conceived and designed the experiments: PSD AEP MHW. Performed the experiments: PSD AEP JRS. Analyzed the data: PSD AEP MHW. Wrote the paper: PSD AEP MHW.