About the Authors

Anna Sandebring

Contributed equally to this work with: Anna Sandebring, Kelly Jean Thomas

Affiliations Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America, Alzheimer's Disease Research Center, Karolinska Institutet, Stockholm, Sweden

Kelly Jean Thomas

Contributed equally to this work with: Anna Sandebring, Kelly Jean Thomas

Affiliation Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America

Alexandra Beilina

Affiliation Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America

Marcel van der Brug

Current address: Department of Molecular and Integrative Neurosciences, The Scripps Research Institute, Jupiter, Florida, United States of America

Affiliation Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America

Megan M. Cleland

Affiliation Surgical Neurology Branch, National Institute of Neurological Diseases and Stroke, Bethesda, Maryland, United States of America

Rili Ahmad

Current address: AstraZeneca R&D Södertälje, Local Discovery RA CNS & Pain Control Disease Biology, Södertälje, Sweden

Affiliation Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America

David W. Miller

Affiliation Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America

Ibardo Zambrano

Affiliation Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America

Richard F. Cowburn

Current address: AstraZeneca R&D Södertälje, Local Discovery RA CNS & Pain Control Disease Biology, Södertälje, Sweden

Affiliation Alzheimer's Disease Research Center, Karolinska Institutet, Stockholm, Sweden

Homira Behbahani

Affiliation Alzheimer's Disease Research Center, Karolinska Institutet, Stockholm, Sweden

Angel Cedazo-Mínguez

Affiliation Alzheimer's Disease Research Center, Karolinska Institutet, Stockholm, Sweden

Mark R. Cookson

Cookson@mail.nih.gov

Affiliation Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AS KJT MvdB RC HB ACM MRC. Performed the experiments: AS KJT AB MvdB MC RA DM IZ. Analyzed the data: AS KJT DM HB ACM MRC. Contributed reagents/materials/analysis tools: MvdB MC HB ACM. Wrote the paper: AS KJT MRC.