About the Authors

David Schley

David.Schley@bbsrc.ac.uk

Affiliation Institute for Animal Health, Pirbright Laboratory, Surrey, United Kingdom

Simon Gubbins

Affiliation Institute for Animal Health, Pirbright Laboratory, Surrey, United Kingdom

David J. Paton

Affiliation Institute for Animal Health, Pirbright Laboratory, Surrey, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DS DJP. Performed the experiments: DS. Analyzed the data: DS SG. Contributed reagents/materials/analysis tools: SG. Wrote the paper: DS SG DJP.