About the Authors

Steven C. Pino

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Bryan O'Sullivan-Murphy

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Erich A. Lidstone

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Chaoxing Yang

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Kathryn L. Lipson

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Agata Jurczyk

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Philip diIorio

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Michael A. Brehm

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

John P. Mordes

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Dale L. Greiner

Affiliations Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America, Molecular Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Aldo A. Rossini

Affiliations Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America, Molecular Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Rita Bortell

rita.bortell@umassmed.edu

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SCP BOM EAL CY AJ Pd DLG RB. Performed the experiments: SCP BOM EAL CY AJ. Analyzed the data: SCP KLL Pd MAB JPM DLG AAR RB. Contributed reagents/materials/analysis tools: JPM. Wrote the paper: SCP KLL MAB JPM DLG AAR RB.