About the Authors

Richard C. Grandison

Contributed equally to this work with: Richard C. Grandison, Richard Wong

Affiliation UCL Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom

Richard Wong

Contributed equally to this work with: Richard C. Grandison, Richard Wong

Affiliation UCL Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom

Timothy M. Bass

Affiliation UCL Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom

Linda Partridge

l.partridge@ucl.ac.uk

Affiliation UCL Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom

Matthew D. W. Piper

Affiliation UCL Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RCG RW TMB LP MDWP. Performed the experiments: RCG RW TMB MDWP. Analyzed the data: MDWP. Contributed reagents/materials/analysis tools: RCG RW TMB MDWP. Wrote the paper: LP MDWP.