About the Authors

Jae Kwak

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Alan Willse

Current address: Monsanto Company, Ankeny, Iowa, United States of America

Affiliation Battelle - Pacific Northwest Division, Richland, Washington, United States of America

Koichi Matsumura

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Maryanne Curran Opiekun

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Weiguang Yi

Current address: Department of Horticulture, University of Georgia, Athens, Georgia, United States of America

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

George Preti

Affiliations Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America, Department of Dermatology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Kunio Yamazaki

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Gary K. Beauchamp

beauchamp@monell.org

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KY GKB. Performed the experiments: KM MCO WY. Analyzed the data: JK AW GP. Contributed reagents/materials/analysis tools: JK AW MCO GP KY. Wrote the paper: JK AW GP KY GKB.