About the Authors

Eric M. Nicholson

Contributed equally to this work with: Eric M. Nicholson, Brian W. Brunelle

Eric.Nicholson@ars.usda.gov

Affiliation Virus and Prion Diseases of Livestock Research Unit, National Animal Disease Center, USDA, Agricultural Research Service, Ames, Iowa, United States of America

Brian W. Brunelle

Contributed equally to this work with: Eric M. Nicholson, Brian W. Brunelle

Affiliation Pre-Harvest Food Safety and Enteric Diseases Research Unit, National Animal Disease Center, USDA, Agricultural Research Service, Ames, Iowa, United States of America

Juergen A. Richt

Affiliation Virus and Prion Diseases of Livestock Research Unit, National Animal Disease Center, USDA, Agricultural Research Service, Ames, Iowa, United States of America

Marcus E. Kehrli Jr

Affiliation Virus and Prion Diseases of Livestock Research Unit, National Animal Disease Center, USDA, Agricultural Research Service, Ames, Iowa, United States of America

Justin J. Greenlee

Affiliation Virus and Prion Diseases of Livestock Research Unit, National Animal Disease Center, USDA, Agricultural Research Service, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EMN BWB JJG. Performed the experiments: EMN BWB. Analyzed the data: EMN BWB. Contributed reagents/materials/analysis tools: JR MEK. Wrote the paper: EMN.