About the Authors

Angela M. Verdoni

Affiliation Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Richard S. Smith

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Akihiro Ikeda

aikeda@wisc.edu

Affiliation Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Sakae Ikeda

aikeda@wisc.edu

Affiliation Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AMV RSS AI SI. Performed the experiments: AMV SI. Analyzed the data: AMV AI SI. Contributed reagents/materials/analysis tools: AMV RSS AI SI. Wrote the paper: AMV AI SI.