About the Authors

Steven M. Woolley

To whom correspondence should be addressed. E-mail: stevenwoolley@wustl.edu

Affiliation Computational Biology Program, Washington University School of Medicine, St. Louis, Missouri, United States of America

David Posada

Affiliation Departamento de Bioquímica, Genética e Inmunología, Facultad de Ciencias, Universidad de Vigo, Vigo, Spain

Keith A. Crandall

Affiliation Department of Biology, Brigham Young University, Provo, Utah, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KC DP SW. Performed the experiments: SW. Analyzed the data: SW. Contributed reagents/materials/analysis tools: SW. Wrote the paper: KC DP SW.