About the Authors

Geraldine A. Wright

*E-mail: jeri.wright@ncl.ac.uk

Affiliation Biology, Newcastle University, Newcastle upon Tyne, United Kingdom

Sonya M. Kottcamp

Affiliation Rothenbuhler Honeybee Laboratory, Ohio State University, Columbus, Ohio, United States of America

Mitchell G. A. Thomson

Affiliation Biology, Newcastle University, Newcastle upon Tyne, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GW SK. Performed the experiments: SK. Analyzed the data: GW MT. Contributed reagents/materials/analysis tools: MT. Wrote the paper: GW MT.