About the Authors

Hiroshi Akashi

To whom correspondence should be addressed. E-mail: akashi@psu.edu

Affiliation Institute of Molecular Evolutionary Genetics, Department of Biology, Pennsylvania State University, State College, Pennsylvania, United States of America

Piyush Goel

Affiliation Institute of Molecular Evolutionary Genetics, Department of Biology, Pennsylvania State University, State College, Pennsylvania, United States of America

Anoop John

Affiliation Institute of Molecular Evolutionary Genetics, Department of Biology, Pennsylvania State University, State College, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HA PG. Analyzed the data: HA PG. Wrote the paper: HA. Other: Developed the C programs: AJ PG HA.