About the Authors

Christian B. Willberg

Affiliations Nuffield Department of Medicine, Oxford University, Oxford, United Kingdom, Division of Experimental Medicine, University of California at San Francisco, San Francisco, California, United States of America

Scott M. Ward

Affiliations Nuffield Department of Medicine, Oxford University, Oxford, United Kingdom, Princess Alexandra Hospital, Woolloongabba, Australia

Reginald F. Clayton

Affiliation Medical Research Council Virology Unit, Glasgow, United Kingdom

Nikolai V. Naoumov

Affiliation The University College London Institute of Hepatology, University College London, London, United Kingdom

Christopher McCormick

Affiliations Faculty of Biological Sciences, Institute of Molecular and Cellular Biology, University of Leeds, Leeds, United Kingdom, Molecular Microbiology and Infection, University of Southampton, Southampton General Hospital, Southampton, United Kingdom

Sandra Proto

Affiliation The University College London Institute of Hepatology, University College London, London, United Kingdom

Mark Harris

Affiliation Faculty of Biological Sciences, Institute of Molecular and Cellular Biology, University of Leeds, Leeds, United Kingdom

Arvind H. Patel

Affiliation Medical Research Council Virology Unit, Glasgow, United Kingdom

Paul Klenerman

To whom correspondence should be addressed. E-mail: paul.klenerman@medawar.ox.ac.uk

Affiliation Nuffield Department of Medicine, Oxford University, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CW PK. Performed the experiments: NN CW SW SP. Analyzed the data: NN CW PK SW. Contributed reagents/materials/analysis tools: NN AP CW PK RC CM MH. Wrote the paper: CW PK SW.