About the Authors

Joy G. Ghosh

Affiliation Department of Biological Structure, University of Washington, Seattle, Washington, United States of America

Scott A. Houck

Affiliation Department of Biological Structure, University of Washington, Seattle, Washington, United States of America

John I. Clark

To whom correspondence should be addressed. E-mail: clarkji@u.washington.edu

Affiliations Department of Biological Structure, University of Washington, Seattle, Washington, United States of America, Department of Ophthalmology, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JG. Performed the experiments: JG SH. Analyzed the data: JC JG SH. Contributed reagents/materials/analysis tools: JG. Wrote the paper: JC JG SH.