About the Authors

Pia Leppänen

Affiliation Department of Biotechnology and Molecular Medicine, A.I. Virtanen Institute, University of Kuopio, Kuopio, Finland

Ivana Kholová

Affiliation Department of Biotechnology and Molecular Medicine, A.I. Virtanen Institute, University of Kuopio, Kuopio, Finland

Anssi J. Mähönen

Affiliation Department of Biotechnology and Molecular Medicine, A.I. Virtanen Institute, University of Kuopio, Kuopio, Finland

Kari Airenne

Affiliation Department of Biotechnology and Molecular Medicine, A.I. Virtanen Institute, University of Kuopio, Kuopio, Finland

Suvi Koota

Affiliation Department of Biotechnology and Molecular Medicine, A.I. Virtanen Institute, University of Kuopio, Kuopio, Finland

Hannu Mansukoski

Affiliation Technical University of Dresden, Dresden, Germany

Johanna Närväinen

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Maria Wirzenius

Affiliation Molecular/Cancer Biology Laboratory, Biomedicum, University of Helsinki, Helsinki, Finland

Leena Alhonen

Affiliation Department of Biotechnology and Molecular Medicine, A.I. Virtanen Institute, University of Kuopio, Kuopio, Finland

Juhani Jänne

Affiliation Department of Biotechnology and Molecular Medicine, A.I. Virtanen Institute, University of Kuopio, Kuopio, Finland

Kari Alitalo

Affiliation Molecular/Cancer Biology Laboratory, Biomedicum, University of Helsinki, Helsinki, Finland

Seppo Ylä-Herttuala

To whom correspondence should be addressed. E-mail: Seppo.Ylaherttuala@uku.fi

Affiliations Department of Biotechnology and Molecular Medicine, A.I. Virtanen Institute, University of Kuopio, Kuopio, Finland, Department of Medicine and the Gene Therapy Unit, Kuopio University Hospital, Kuopio, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SY PL KA JJ. Performed the experiments: IK AM SK HM JN. Analyzed the data: SY PL IK AM JN. Contributed reagents/materials/analysis tools: KA MW LA. Wrote the paper: SY PL IK AM KA