A ‘Terror of Tyrannosaurs’: The First Trackways of Tyrannosaurids and Evidence of Gregariousness and Pathology in Tyrannosauridae

• Richard T. McCrea ,

  * E-mail: rtmccrea@prprc.com

  Affiliations Peace Region Palaeontology Research Centre, Tumbler Ridge, British Columbia, Canada, Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Alberta, Canada

  ⨯
• Lisa G. Buckley,

  Affiliations Peace Region Palaeontology Research Centre, Tumbler Ridge, British Columbia, Canada, Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

  ⨯
• James O. Farlow,

  Affiliation Department of Geosciences, Indiana-Purdue University, Fort Wayne, Indiana, United States of America

  ⨯
• Martin G. Lockley,

  Affiliation Dinosaur Trackers Research Group, University of Colorado at Denver, Denver, Colorado, United States of America

  ⨯
• Philip J. Currie,

  Affiliation Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

  ⨯
• Neffra A. Matthews,

  Affiliation Bureau of Land Management, Denver, Colorado, United States of America

  ⨯
• S. George Pemberton

  Affiliation Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Alberta, Canada

  ⨯