Referee 2's review (Christian R. Voolstra):
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N.B. These are the comments made by the referee when reviewing an earlier version of this paper. Prior to publication, the manuscript has been revised in light of these comments and to address other editorial requirements.
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Thorough study, concise written, important implications;
Would like to see more discussion on possible role of parrotfish (reef animals in general) on dispersal of Symbiodinium spp. and how an experiment would look like that proves a role of fish in dispersal of zooxanthellae;
please also discuss: if Symbiodinium spp. live within bedforming macroalgae how do they transfer back to coral?